Yves Alarie

Yves Alarie

Full Professor

Department of Biology
Science, Engineering and Architecture
S-623, Science Building Sudbury Campus


Born in Trois-Rivières, Qc. I received a college degree in Health Sciences from CEGEP de Trois-Rivières in 1976; B.A. Philosophy and Anthropology (Université laval in 1980), BSc and MSc in Bio-Ecology from Université du Québec à Trois-Rivières in 1983 and 1985 respectively; PhD Sciences biologiques (Université de Montréal, 1990); NSERC Postdoctoral fellow (University of Manitoba (1990-1992). Assistant Professor University of Manitoba 1992-93 and Laurentian University 1993-1995. I have been promoted to Full professor  July 1st 2000.My research program is in the field of water beetle systematics; I have received the Laurentian University Research Excellence Award in 2002-2003. Director of RDC at Laurentian since October 2011. Married, father of 3 children.


  • PhD Sciences biologiques (Entomologie) (Université de Montréal)
  • MSc Ecologie (Université du Québec à Trois-Rivières)
  • BSc Bio-Ecologie (Université du Québec à Trois-Rivières)
  • B. A. Philosophie et Anthropologie (Université Laval)
  • DEC Sciences Santé (CEGEP de Trois-Rivières)

Academic Appointments

Director of Research Development and Creativity at Laurentian 2011-2015.


My research emphasizes study of larval morphology of aquatic beetles. Although water beetles are among the most common insect inhabitants of freshwaters, knowledge of their larval morphology is scanty throughout the world. The identification of larvae is a continuing problem because the literature available to accomplish this is scattered, limited to certain groups, outdated, difficult to use or non-existent. Because of this, I developed a framework, which facilitates comparison among species. My system of analysis focuses on study of body sensilla, which are useful both for diagnosis and study of the genealogical relationships among taxa. The study of larval features should allow more comprehensive classifications to be constructed. Larvae are under differing selection pressures and therefore show quite different features. A long-term goal of my research is to produce a more rigorous and stable classification by combining characters from larvae with other data sets including adult and molecular characters.


  • Laurentian University Research Excellence Award 2002-2003
  • NSERC Postdoctoral Fellow 1990-1992 (University of Manitoba, Winnipeg, Manitoba)


General Entomology (3cr.)

Medical Entomology (3cr.)

Entomologie Générale (3 cr.)

Entomologie Médicale (3 cr.)

Méthodes Expérimentales en Biologie (3 cr.)


132. Degasparro, S. L., Alarie,Y. and Beresford D. . 2020. Faunistic survey of Dytiscidae (Coleoptera: Adephaga) from Ontario's Far North. The Coleopterists Bulletin (submitted).

131. Gustafson, G., Miller, K. B., Michat, M. C., Alarie, Y., Bacca, S. and Short, A.E. Z. 2020. The enduring value of reciprocal illumination in the era of insect phylogenomics: a response to Cai et al. 2020. Systematic Entomology (submitted).

130. Urcola, J.I., Alarie, Y., Benetti, C.J. and Michat, M.C. 2020. Description of the larval stages of Hydrocanthus socius Sahlberg, 1844 (Coleoptera: Noteridae) with chaetotaxy analysis. Annales Zoologici (in press).

129. Michat, M. C., Archangelsky, M. and Alarie, Y.  2020. Morphology and chaetotaxy of Neotropical Haliplus larvae (Coleoptera: Haliplidae). Revista Mexicana de Biodiversidad, (in press).

128. Okada, R. and Alarie Y. 2020. Description of the immature stages of the  endangered Japanese endemic Oreodytes kanoi (Kamiya, 1938) (Coleoptera: Dytiscidae, Hydroporinae) and comparison with the known larvae of Oreodytes Seidlizt, 1887. Zootaxa 4820(1): 001-018.

127. Urcola, J.I., Benetti, C.J., Alarie, Y.Rodriguez, G. and Michat, M.C. 2020. The  unknown larva of the burrowing water beetle genus Liocanthydrus Guignot, 1957 (Coleoptera: Noteridae). Journal of Natural History (submitted).

126. Urcola, J.I., Benetti, C.J., Alarie, Y., Rodriguez, G. and Michat, M.C. 2020. Characterization and mapping of sensilla on the head appendages of noterid larvae (Coleoptera: Noteridae), and development of a preliminary biometric method for taxa delimitation Journal of morphology, 281: 1210- 1222. https://doi.org/10.1002/jmor.21241.

125. Alarie, Y. 2020. The Hydradephaga (Coleoptera, Haliplidae, Gyrinidae,  Dytiscidae) fauna of Manitoulin Island, Ontario, Canada, the world largest freshwater island. The Coleopterists Bulletin (in press).

124. Tsyrlin, E., Carew M. and Alarie, Y. 2020. Redescription of larvae of Chostonectes nebulosus (MacLeay, 1871) (Coleoptera: Dytiscidae: Hydroporinae: Hydroporini: Sternopriscina) with a key of identification of the known larvae of Chostonectes Sharp 1882. Zootaxa 4718(3): 436-446. 

123. Alarie, Y., M.C. Michat and Watts C.H.S. 2020. Larval morphology of Megaporus Brinck, 1943 (Coleoptera: Dytiscidae): description of M. hamatus (Clark, 1862) and M. gardnerii (Clark, 1862) and phylogenetic considerations. The Coleopterists Bulletin 74(1): 139-160.

122. Alarie, Y. and M. C. Michat. 2020. Larval morphology of Agabinae (Coleoptera: Dytiscidae): descriptions of three species of the subgenus Agabus s. str. Leach,  1817 with phylogenetic considerations. Aquatic Insects: 41(3): 197-226. https://doi.org/10.1080/01650424.2020.1741646

121. Michat, M. C., Alarie, Y., and Watts C.H.S 2019. Dealing with a hairy beast – Larval morphology and chaetotaxy of the Australian endemic diving-beetle genus Spencerhydrus Sharp, 1882 (Coleoptera: Dytiscidae, Cybistrini). Zookeys 884: 53-67.

120. Urcola, J.I., Alarie, Y.Benetti, C.J. and Michat, M.C. 2019. Larval morphology of Suphisellus Crotch, 1873 (Coleoptera: Noteridae): description of first instarof S. rufipes (Sharp, 1882) with biological notes and chaetotaxy analysis. Annales Zoologici 69(4): 817-825.

119. Hajek, J., Alarie, Y. Stastny, J. and Vondracek. 2019. The first hygropetric Platynectes and its larva from Eastern China (Coleoptera: Dytiscidae). Acta Entomologica Musei Nationalis Pragae, 59(1): 217-228.

118. Alarie, Y. 2019. Further contributions to the Hydradephaga (Coleoptera, Haliplidae, Gyrinidae, Dytiscidae) fauna of Cape Breton Island, Canada: new records, distributions and faunal composition. ZooKeys 897: 49-66.

117. Okada, R., Alarie, Y. and Michat, M.C. 2019. Description of the larvae of four Japanese Platambus Thomson, 1859 (Coleoptera: Dytiscidae: Agabinae) with phylogenetic considerations. Zootaxa 4646(3): 401-433.

116. Urcola J.I., Benetti C.J., Alarie, Y., Torres, P.L.M. and Michat, M.C. 2019. Morphology and chaetotaxy of the instars of Hydrocanthus sharpi Zimmermann, 1928 (Coleoptera: Noteridae). The Coleopterists Bulletin 73(3): 611-620.

115. Alarie, Y., M.C. Michat and Watts, C.H.S. 2019. Larval morphology of the  Australian radiation Sternopriscina (Coleoptera: Dytiscidae, Hydroporinae): description and phylogenetic placement of the genus Chostonectes Sharp, 1880.                Aquatic Insects 40(4): 328-354.

114. Vasilikopoulos, A., Balke, Beutel, R. G., Donath, A., Podsiadlowski, L., Pflug, J. M.,Waterhouse,R. M., Meusemann, K., Peters, R. S., Hescalona, H. E., Mayer, C., Liu, S., Hendrich, L., Alarie, Y., Bilton,D. T., Jia, F., Zhou, X., Ribera, I.,  Maddison, D. R., Niehuis, O., and Misof. B. 2019. Phylogenomics of the superfamily Dytiscoidea (Coleoptera: Adephaga) with an empirical evaluation of phylogenetic conflict and systematic error.  Molecular Phylogenetics and  Evolution 135: 270-285.

113. Urcola, J.I., Alarie, Y., Benetti, C.J., Rodriguez, G. and Michat, M.C. 2019. Larval morphology and analysis of primary chaetotaxy in the genus Suphis (Coleoptera: Noteridae). Zootaxa, 4619(1): 121-138.

112. Alarie, Y., Michat, M.C. Jia, and Hajek, J., F. 2019. Hydrotrupes chinensis Nilsson, 2003 (Coleoptera: Dytiscidae: Agabinae): new records, description of  larvae, and notes on its biology and phylogenetic relationships.  Aquatic Insects 40(3): 236-256.

111. Benetti, C.J., Michat, M.C., Alarie, Y. and Hamada, N. 2019 Description of the second and third instars of Platynectes decemnotatus (Aubé, 1838) (Coleoptera: Dytiscidae). Zootaxa 4544(3): 381-394.

110. DeGasparro, S.D., Brown, G., Alarie, Y. and Beresford, D.V. 2018. Predaceous diving beetle records form Akimiski Island, Nunavut (Coleoptera: Dytiscidae). The Coleopterists Bulletin 72(4): 866-869.

109.  Michat, M.C., Alarie, Y. and Hajek, J. 2018. Larvae of Huxelhydrus syntheticus Sharp, 1882 (Coleoptera: Dytiscidae: Bidessini). Zootaxa, 4524(1): 121-131.

108.  Alarie, Y., M.C. Michat and Watts C.H.S. 2018. Larval morphology of Barretthydrus (Coleoptera: Dytiscidae): descriptions of B. gemminatus and B.tibialis and phylogenetic considerations. The Coleopterists Bulletin 72(4): 639-661.

107.  Michat, M.C., Alarie, Y. and Miller, K. B. 2017. Higher-level phylogeny of diving beetles (Coleoptera: Dytiscidae) based on larval characters. Systematic Entomology, 42: 734-767.

106.  Alarie, Y., Michat, M.C. and G. L. Challet. 2017. Larval Description and Phylogenetic Placement of the South Africa Endemic genera Coelhydrus and Darwinidrus (Coleoptera: Dytiscidae, Hydroporinae, Hyphydrini). The Colepterists Bulletin 69(3): 489-497.

105. Ashbee, H., Marshall, S. and Alarie, Y. 2017. Haliplidae (Coleoptera) of Eastern  Canada. Journal of Arthropods Identification,  doi:10.3752/cjai.2017.32 

104. M. Balke, Alarie, Y., and Beutel, R.. 2016.  Meruidae. In Handbook of Zoology/Handbuch der Zoologie. Ed. by Beutel, Rolf G. / Kristensen, Niels Peder / Leschen, Richard / Purschke, Günter / Westheide, Wilfried / Zachos, Frank

103. Vondel, B. J. van and Alarie Y. 2016. A New Species of Haliplus Latreille, 1802 (Coleoptera: Adephaga: Haliplidae) from Canada. The Coleopterists Bulletin 70(4): 801-804.

102. Alarie, Y. 2016. Further contributions to the Hydradephaga (Coleoptera, Haliplidae, Gyrinidae, Dytiscidae) fauna of Prince Edward Island, Canada: new records, distributions and faunal composition. Zookeys, 600: 103-129.

101. Michat, M.C., T. M. Alvarenga, M. S. Silva, Y. Alarie. 2016. First larval description and chaetotaxic analysis of the neotropical whirligig beetle genus Enhydrus Laporte, (Coleoptera: Gyrinidae). Revista Brasileira de Entomologia, 60: 231-237.

100. Michat, M.C., Alarie, Y. and Watts, C.H.S. 2015. Phylogenetic Placement of the Genus Sternhydrus (Coleoptera: Dytiscidae: Cybistrini) Based on Larval Morphology, With Description of S. atratus. Annals of Entomological Society of                                                                                            America, 108(5): 881-892

99. Michat, M.C., Alarie, Y. and Hajek, J. 2015. Larval morphology of Yola Gozis, 1886 (Coleoptera: Dytiscinae: Bidessini) and comparison with other genera of the tribe Bidessini. The Coleopterists Bulletin 69(3): 489-497.

98. Alarie, Y. and M. C. Michat. 2014. Bridging Ecology and Systematics: 25 years of study of larval morphology of World Dytiscidae, 17-47. In Ecology, Systematics, and the Natural History of Predaceous Diving Beetles (Coleoptera:                Dytiscidae). (D.A. Yee, editor). Springer, The Netherlands, xvii + 484 pp.

97. Michat, M.C., Alarie, Y., Fenglong, J., Shengquan, W., Hájek, J. and Balke, M. 2014. Description of the second- and third instars of Aspidytes wrasei Balke, Ribera & Beutel, 2003 (Coleoptera: Aspidytidae), with a key for the identification of larvae of Aspidytes Ribera, Beutel, Balke & Vogler, 2002 and  phylogenetic considerations. Zootaxa 3881 (4): 362-372.

96. Michat, M.C., Alarie, Y. and Hendrich, L. 2014. Description of the third instar of Hygrobia nigra(Clark) (Coleoptera: Hygrobiidae), with a key for identification of mature larvae of Hygrobia Latreille and phylogenetic analysis.  Zootaxa                        3827(3): 318-330.

 95. Barman, E.H., Michat, M.C., Y. Alarie, and Wolfe, G.W. 2014. Description of the first-instar larva of  Hoperius planatus, with phylogenetic implications. The Coleopterists Bulletin. 68(2): 321-330.

94. Miller, K.B. Jean, A., Alarie, Y., Hardy, N. and GibsonJ.R. 2013. Phylogenetic Placement of North American Subterranean Diving Beetles (Coleopera: Dytiscidae), Arthropods Systematics and Phylogeny, 71(2): 75-90.

93. Alarie, Y., J.R. Gibson and Miller, K.B. 2013. Descriptions of Larvae of the North American Endemic Stygobiontic Ereboporus naturaconservatus Miller, Gibson and Alarie and Haideoporus texanus Young and Longley (Coleoptera:                               Dytiscidae). Tijdschrift voor Entomologie, 156: 1-10.

 92. Michat, M.C. and Alarie, Y. 2013. Phylogenetic relationships, larval morphology and chaetotaxy of the subfamily Coptotominae (Coleoptera: Dytiscidae), The Canadian Entomologist, 145: 247-264.

91. Michat, M.C., Alarie, Y. and Watts. C.H.S. 2012. Description of the larvae of 30 species of Limbodessus Guignot (Coleoptera: Dytiscidae: Bidessini), ground plan of chaetotaxy of the genus, comparative morphology of epigean and                           stygobitic phenotypes, and phylogenetic analysis. Zootaxa 3584: 1–110.

90. Short, A.E.Z., Alarie, Y., Garcia, M. and L.J. Joly. 2012. Are noterids specialized meruids (Coleoptera, Adephaga)? A reply to Dressler et al. Systematic Entomology 37:417-419.

89. Alarie, Y. and D.T. Bilton. 2012. The larva of Hydroporus zimmermanni G. Müller, 1926 (Coleoptera: Dytiscidae: Hydroporinae), with notes on its ecology and a review of described Hydroporuslarvae. The Coleopterists Bulletin, 66(2):                81-92.

88. Alarie, Y., Michat, M.C. and K.B. Miller. 2011. Notation of primary setae and pores on larvae of Dytiscinae (Coleoptera: Dytiscidae), with phylogenetic considerations. Zootaxa 3087: 1-55.

87. Toledo, M., Megna, Y. S., and Y. Alarie. 2011. Description of a new species of Laccodytes  Régimbart, 1895 (Coleoptera, Dytiscidae, Laccophilinae) from Cuba.  Zootaxa 2792: 63-67.

86. Alarie, Y., Short, A. E. Z., M. Garcia, and L. Joly. 2011. Larval Morphology of  Meruidae and its phylogenetic implications (Coleoptera: Hydradephaga). Annals of the Entomological Society of America 104(1): 25-36.

85. Michat, M.C., Alarie, Y. and Watts. C.H.S. 2011. Larval morphology of Allodessus Guignot (Coleoptera: Dytiscidae). Aquatic Insects 33: 27-40.

84. Balke, M., Wewalka, G., Alarie, Y. and Ribera, I. 2010. Dytiscidae: the genus Rhantus Dejean. In Water Beetles of New Caledonia, 129-148. Jach, M. &  Balke, M. (eds.) Koleopterologische Rundschau 459 pp.

83. Michat, M.C., Alarie, Y. and Watts, C.H..S. 2010. Descriptions of the first instar larva of the hypogaeic species Neobidessodes limestoneensis (Watts & Humphreys) and of the third instar larva of Hydroglyphus balkei Hendrich                         (Coleoptera: Dytiscidae: Bidessini) with phylogenetic considerations. Zootaxa 2658: 38-50.

82. Alarie, Y. and Longing, S. 2010. Description of the larvae of Heterosternuta sulphuria (Matta and Wolfe) with a key of identification of the known species of Heterosternuta Strand. The Coleopterists Bulletin  64: 85-90.

81. Alarie, Y., Michat, M.C. and C.H.S. Watts. 2009. Larval morphology of Paroster  Sharp, 1882 (Coleoptera: Dytiscidae: Hydroporinae): reinforcement of the hypothesis of monophyletic origin and discussion of phenotypic accommodations to a subterranean environment. Zootaxa 2274: 1-44.

80. Michat, M. C. and Alarie, Y. 2009. Phylogenetic relationships of Notaticus  Zimmermann (Coleoptera: Dytiscidae) based on larval morphology. Annals of the Entomological Society of America102(5): 797-808.

79. Miller, K.B., Gibson, J.R. and Alarie, Y. 2009. North American stygobiontic diving beetles (Coleoptera: Dytiscidae: Hydroporinae) with description of Ereboporus naturaconservatus Miller, Gibson and Alarie, new genus and                          species from Texas, USA. The Coleopterists Bulletin 63(2): 191-202.

78. Alarie, Y., Michat, M.C., Nilsson, A.N., Archangelsky, M. and L. Hendrich  2009. Larval morphology of Rhantus Dejean (Coleoptera: Dytiscidae: Colymbetinae): descriptions of 21 species and phylogenetic considerations.  Zootaxa, 2317: 1-102.

77. Alarie, Y., Megna, Y. S.and A. Deler-Hernández. 2009.  First West Indies records of Thermonectus succinctus (Aubé, 1838), with notes on other Cuban species  (Coleoptera: Dytiscidae). Koleopterologische Rundschau 79: 5-16.

76. Fairn, E., Alarie, Y. and A. Schulte-Hostedde. 2009. Temporal variation of adult sex ratio in the whirligig beetle Dineutus nigrior Roberts (Coleoptera: Gyrinidae). The Coleopterists Bulletin, 63(3): 298-300.

75. Alarie, Y.  2009. Hydradephaga of the Magdalen Island Archipelago. Koleopterologische Rundschau 79: 17-25.

74. Fairn, E. R., A. I. Schulte-Hostedde, and Y. Alarie. 2008. Attachment site and  infestation parameters of parasitic water mites on the whirligig beetle Dineutus nigrior Roberts (Coleoptera: Gyrinidae). (Coleoptera: Gyrinidae). Aquatic                         Insects 30(3): 223-228.

73. Fenske-Babin, J., Anand, M. and Y. Alarie. 2008. Rapid morphological change in whirligig beetle (Coleoptera: Gyrinidae) museum species correlate with climate change. Ecological Entomology 33: 646-651.

72. Fairn, E., Schulte-Hostedde, A. and Y. Alarie. 2008. Water mite parasitism is associated with body condition and sex of the whirligig beetle Dineutus nigrior Roberts (Coleoptera: Gyrinidae). Ecoscience 15(3): 327-331.

71. Michat, M.  and Y. Alarie.  2008. Morphology and chaetotaxy of larval Hypodessus cruciatus (Régimbart) (Coleoptera: Dytiscidae: Hydroporinae), and analysis of the phylogenetic relationships of the Bidessini based on larval characters. Studies on Neotropical Fauna and Environment 43(2): 135-146.

70. Fairn, E., Alarie, Y. and A. Schulte-Hostedde. 2007. Sexual size dimorphism in the diving beetle Laccophilus maculosus Say (Coleoptera: Dytiscidae). The Coleopterists Bulletin 61: 409-418.

69. Miller, K.B., Alarie, Y. and Whiting, M. F. 2007. Description of the Larva of Notaticus fasciatus Zimmermann (Coleoptera: Dytiscidae ) associated with adults using DNA sequence data.  Annals of the Entomological Society of America 100: 787-797.

68. Alarie, Y. and M. Michat. 2007. Phylogenetic analysis of Hydroporinae (Coleoptera:Dytiscidae) based on larval morphology, with description of first-instar Laccornellus  lugubrisAnnals of the Entomological Society of America 100: 655-665.

67. Fery, H., Petrov, P.N. and Alarie, Y.  2007.  Notes on the identity of Oerodytes dauricus (Motschulsky, 1860) and other members of the O. alaskanus – clade (Coleoptera: Dytiscidae). Linzer Biologische  Beiträge 39: 65-78.

66. Michat, M. C.  Alarie, Y., P.L.M. Torres and Y. Suarez. 2007. Larval morphology of the diving beetle Celina and the phylogeny of ancestral hydroporines. Invertebrate Systematics 21: 239-254.

65. Alarie, Y., M. C. Michat, Archangelsky, M., and H. M. James. 2007. Larval morphology of Liodessus Guignot: generic characteristics, descriptions of five species and comparisons with other members of the tribe Bidessini (Coleoptera:                Dytiscidae: Hydroporinae).  Zootaxa 1516: 1-21.

64. Alarie, Y. and M. C. Michat.  2007. Primary setae and pores on the maxilla of larvae of the subfamily Hydroporinae (Coleoptera: Adephaga: Dytiscidae): ground plan pattern reconsidered. The Coleopterists Bulletin 61: 310-317.

63. Fairn, E., Alarie, Y. and A. Schulte-Hostedde. 2007. Sexual dimorphism in Dineutus nigrior Roberts Coleoptera: Gyrinidae). The Coleopterists Bulletin 61: 113-120.

62. Fairn, E., A. Schulte-Hostedde and Alarie, Y. 2007. Allometry and sexual selection in the whirligig beetle Dineutus nigrior (Coleoptera: Gyrinidae). Ethology 113: 257-266.

61. Balke, M., Wewalka, G., Alarie, Y. and Ribera, I. 2007. Molecular Phylogeny of Pacific Colymbetini: radiation of New Caledonian and Fiijan species. Zoologica Scripta  36: 173-200.

60. Alarie, Y.  2006. Amphizoidae, Aspidytidae, Haliplidae, Noteridae and Paelobiidae (Coleoptera, Adephaga). World Catalogue of Insects, Volume 7. The Coleopterists Bulletin 60: 305-306

59. Michat, M. and Alarie, Y. 2006. The larvae of Amarodytes duponti (Aubé) (Coleoptera: Dytiscidae: Hydroporinae), with comments on Bidessini larval morphology and chaetotaxy.  Zootaxa 1351: 1-13.

58. Cook, L. Alarie, Y., and Babin-Fenske, J. 2006. Morphometric Analysis of Dineutus emarginatus  Say (Coleoptera: Gyrinidae); subspecies designation reconsidered. The Coleopterists Bulletin 60: 235-241.

57. Arnott, S.E, Jackson, A., and Y. Alarie2006. The Distribution and Potential Effects of Water Beetles in Lakes Recovering from Acidification.  Journal of the North American Benthological Society 25: 811-824.

56. Alarie, Y. and G. L. Challet. 2006.  Larval description and phylogenetic placement of the South African endemic genus Andex Sharp (Coleoptera: Adephaga: Dytiscidae).  Annals of the Entomological Society of America  99: 743-754.

55. Alarie, Y. and Y. S. Megna. 2006. Larval morphology of Cuban specimens of Pachydrus obniger (Chevrolat)  (Coleoptera: Adephaga: Dytiscidae). Koleopterologische Rundschau 76: 43-49.

54. Shaverdo, H. and Y. Alarie. 2006. Description of larvae of Canthyporus kenyensis Bilardo & Sanfilippo (Coleoptera: Dytiscidae: Hydroporinae) with implication for the phylogeny of the tribe Hydroporini. Aquatic Insects. 28(2): 113-130.

53. Schulte-Hostedde, A. and Y. Alarie.  2006. Morphological patterns of sexual selection in the diving beetle Graphoderus liberusEvolutionary Ecology Research 8: 891-901.

52. Belzile, N., Chen, Y-W. Gunn. J. M., Tong, J., Alarie, Y., Delonchamp, T. and C- Y. Yang. 2006. The effect of selenium on mercury assimilation by freshwater organisms. Canadian Journal of Fisheries and Aquatic Sciences 63: 1-10.

51. Alarie, Y. and S. Hughes. 2006.  Larval morphology of Hoperius Fall and MelademaLaporte  (Coleoptera: Adephaga: Dytiscidae) and phylogenetic implications  for  the tribe Colymbetini. Bollettino  della Societa Entomologica Italiana, Genova. (INVITED PAPER) 85: 307-334.

50. Alarie, Y. and G. L. Challet.  2006. Study of the larvae of Prismopes suturalis  Sharp (Coleoptera: Dytiscidae, Hydroporinae) with implications for the phylogeny of the Hyphydrini. Aquatic Insects 28: 31-46

49. Miller, K.B., Alarie, Y., Wolfe, G.W. and Whiting, M. F. 2005. Association of adults and larvae of Holometabola using DNA sequences: the larvae of Philodytes umbrinus(Motschulsky) (Coleoptera: Dytiscidae). Systematic Entomology  30: 499-509

48. Alarie, Y. and D. T. Bilton. 2005.  Larval morphology of the new adephagan family Aspidytidae (Coleoptera: Adephaga) with phylogenetic implications. Annals of the  Entomological Society of America 98(4): 417-430.

47. Alarie, Y. and C.H.S. Watts. 2005.  Larval morphology of four species of the Hyphydrus lyratus species-group (Coleoptera: Dytiscidae: Hydroporinae). Australian Journal of Entomology 44: 244-251.

46. Alarie, Y. 2005. World Catalogue of Insects, volume 3, Dytiscidae (Coleoptera). Annals of the Entomological Society of America 98: 150.

45. Alarie, Y. and L.-J. Wang.  2004.  Larval morphology of the Taiwan endemic species Rhantus formosanus Kamiya (Coleoptera: Adephaga: Dytiscidae). Koleopterologische Rundschau 74: 147-156

44. Alarie, Y., R.G. Beutel, and C.H.S. Watts.  2004.  Larval morphology of the Hygrobiidae (Coleoptera: Adephaga: Dytiscoidea) with phylogenetic considerations. European Journal of Entomology  101:  293-311.

43. Babin. J. and Y. Alarie.  2004.  Taxonomic revision of genus Gyretes Brullé (Coleoptera: Gyrinidae) from America north of Mexico. The Coleopterists Bulletin  58: 538-567.

42. Alarie, Y. and C.H.S. Watts.  2004. Larvae of the dytiscid genus Antiporus Sharp, 1882 (Coleoptera: Dytiscidae) and phylogenetic placement in the tribe Hydroporini. Invertebrate Systematics  18: 523-546.

41. Alarie, Y. and C.H.S. Watts 2003.  Description of 1st and 3rd instar of Allomatus nannup Watts (Coleoptera: Adephaga: Dytiscidae).  The Coleopterists Bulletin 57(3): 255-264.

40. Alarie, Y. and F. Butera.  2003.  The larva of Matus leechi Young (Coleoptera: Adephaga: Dytiscidae): implications for the larval groundplan of Matus Aubé. Aquatic Insects 25(1): 63-70.

39. Alarie, Y., P.J. Spangler, and W.E. Steiner.  2002.  Larval morphology of Agabetes Crotch (Coleoptera: Adephaga: Dytiscidae): the hypothesis of sister-group relationship with the subfamily Laccophilinae revisited. The Coleopterists Bulletin 56(4): 547-567.

38. Alarie, Y.M. Archangelsky, A.N. Nilsson and C.H.S. Watts 2002.  Larval  morphology of the genus Lancetes Sharp (Coleoptera: Adephaga) descriptions of six species and phylogenetic relationships. The Canadian Entomologist 134:                 467-501.

37. Alarie, Y., C.H.S. Watts, and A.N. Nilsson.  2001. Study of the larval morphology  of he tribe Matini (Coleoptera: Dytiscidae, Colymbetinae): descriptions of Batrachomatus daemeliMatus bicarinatus, and Allomatus nannup and      phylogenetic relationships. The Canadian Entomologist 133: 165-196.

36. Alarie, Y., Jan G.M. Cuppen, L. Hendrich, and A.N. Nilsson.  2001.  Description  of larvae of Herophydrus musicus (Klug 1833) and analysis of relationships with members of the genus Hygrotus Stephens (Coleoptera: Dytiscidae, Hydroporinae).  Aquatic Insects 23: 193-207.

35. Alarie, Y. and G. Wewalka.  2001. Description of the mature larva of Glareadessus stocki Wewalka and Bistrom (Coleoptera: Dytiscidae), a stygobiontic Bidessini from the Persian Gulf region. The Coleopterists Bulletin 55: 144-151.

34. Alarie, Y., P. Wood, A.M.H. deBruyn and J.G.M. Cuppen.  2001.  Descriptions of larvae of Hydroporus ferrugineus Stephens and H. polaris Fall (Coleoptera: Adephaga: Dytiscidae).  Aquatic Insects 23:123-133.

33. Alarie, Y. and D. T. Bilton.  2001. Larval morphology of Hydrotarsus Falkenström: Generic characteristics, description of H. compunctus (Wollaston 1865), and analysis of relationships with other members of the tribe Hydroporini (Coleoptera Dytiscidae, Hydroporinae). The Coleopterists Bulletin 55: 341-350.

32. Larson, D. J., Y. Alarie and R. E. Roughley. 2000. Predaceous Diving Beetles (Coleoptera:Dytiscidae) of the Nearctic Region, with an emphasis on the fauna of Canada and Alaska.  NRC Research Press, Ottawa, Ontario, Canada, 982 pp.

 31. Alarie, Y., A. N. Nilsson, Hendrich, L. and C. H. S. Watts. 2000.  Larval morphology of four genera of Laccophilinae (Coleoptera: Dytisicidae) with an analysis of their  phylogenetic relationships.  Insect Systematics and Evolution 31: 121-164.

30. Alarie, Y., A.N. Nilsson, and L. Hendrich 1999.  Larval morphology of the Palaearctic genera Deronectes Sharp and Scarodytes Gozis (Coleoptera: Dytiscidae: Hydroporinae) with implications for the phylogeny of the Deronectes-group of genera. Entomologica scandinavica 30: 173-195.

29. Paquette, D. and  Y. Alarie.  1999. Species composition of Dytiscid beetles (Coleoptera: Dytiscidae) in cattail ponds of northeastern Ontario, Canada.  The Coleopterists Bulletin 53(2): 160-166.

 28. Alarie, Y., and J. A. Delgado.  1999.Study of the larvae of Antiporus strigosulus  (Brown) (Coleoptera: Dytiscidae) with implications for the phylogeny of the  Hydroporini.  Aquatic Insects  21(2): 99-113.

27. Alarie, Y., and M. Balke.  1999.  Study of larvae of Carabdytes upin Balke,  Hendrich & Wewalka (Coleoptera: Adephaga: Dytiscidae) with implications for  the phylogeny of the Colymbetinae.  Coleopterists Bulletin 53(2): 146-154.

26. Alarie, Y., and D. J. Larson.  1998.  Description of larvae of Agabinus glabrellus (Coleoptera: Dytiscidae: Colymbetinae).  The Coleopterists Bulletin 52: 339-350.

25. Alarie, Y.  1998.  Primary setae and pores on the cephalic capsule and head appendages of the Colymbetinae (Coleoptera: Adephaga: Dytiscidae).  The Canadian Entomologist 130: 803-824.

24. Alarie, Y., P.J. Spangler,  and P. D. Perkins  1998.  Study of the larvae of Hydrotrupes palpalis Sharp (Coleoptera: Adephaga: Dytiscidae) with  implications for the phylogeny of the Colymbetinae.  The Coleopterists Bulletin 52: 313-332.

23. Alarie, Y.,  H. Joly, and D. Dennie  1998.  Cuticular hydrocarbon analysis of the aquatic beetle Agabus anthracinus Mannerheim (Coleoptera: Dytiscidae).  The Canadian Entomologist 130: 615-629.

22. Alarie, Y., and K. Fritz. 1998. Description of larvae of Heterosternuta diversicornis  (Sharp) (Coleoptera: Dytiscidae: Hydroporinae). Entomologica scandinavica 29: 39-46.

21. Alarie, Y., L.-J. Wang, A.N. Nilsson, and P.J. Spangler 1997. Larval morphology of four genera of the tribe Hyphydrini Sharp (Coleoptera: Dytiscidae, Hydroporinae) with an analysis of their phylogenetic relationships.  Annals of the Entomological Society of America 90: 709-735.

20. Alarie, Y. and  A.N. Nilsson  1997.  Larvae of Stictonectes (Brink): Generic   characteristics, description of S. canariensis Machado, and analysis of relationships with other members of the tribe Hydroporini (Coleoptera: Dytiscidae).  The Coleopterists Bulletin 51(2): 107-121.

19. Alarie, Y.  1997.  Taxonomic Revision and Phylogenetic Analysis of the genus Oreodytes Seidlitz (Coleoptera: Dytiscidae: Hydroporinae) Based on larval morphology. The Canadian Entomologist 129: 399-503.

18. Alarie, Y., L.-J. Wang, and P.-S. Yang. 1996. The description of the larvae of Neonectes babai Satô (Coleoptera; Dytiscidae: Hydroporinae) with a discussion of its phylogenetic relationships with members of the Oreodytes scitulus    species-group. The Coleopterists Bulletin 50(4): 373-381.

17. Alarie, Y. and Nilsson, A.N.  1996.  The larvae of Neonectes J. Balfour-Browne (Coleoptera: Dytiscidae: Hydroporinae) with a description of N. natrix (Sharp 1884) and a discussion of its phylogenetic relationships with members of the  tribe Hydroporini Sharp. The Coleopterists Bulletin  50(2): 45-53.

16. Alarie, Y.  1995.  Description of the larval stages of Hydroporus (s.str.) dentellus  Fall 1917 (Coleoptera: Dytiscidae: Hydroporinae). The Coleopterists  Bulletin  49(2): 157-168.

15. Alarie, Y.  1995.  Primary setae and pores on the legs, the last abdominal segment, and the urogomphi of larval Colymbetinae (Coleoptera: Dytiscidae) wih an analysis of their phylogenetic relationships. The Canadian Entomologist  127: 913-943.

14. Alarie, Y.  1994.  Taxinomie assistée par ordinateur.  Revue d'entomologie du Québec  37: 36-48.

13. Alarie, Y.  1993.  A systematic review of the North American species of the Oreodytes alaskanus clade (Coleoptera: Dytiscidae: Hydroporinae). The Canadian Entomologist 125: 847-867.

12. Alarie, Y. 1992.  Description of the larval stages of Hydroporus (Heterosternutacocheconis Fall 1917 (Coleoptera: Dytiscidae: Hydroporinae) with a key of the known larvae of Heterosternuta Strand. The Canadian Entomologist 124: 827-840.

11. Alarie, Y. 1991.  Primary setae and pores on the cephalic capsule and head appendages of larval Hydroporinae (Coleoptera: Dytiscidae).  Canadian Journal of Zoology  69: 2255-2265.

10. Alarie, Y. 1991.  Description of larvae of 17 Nearctic species of Hydroporus Clairville (Coleoptera: Dytiscidae: Hydroporinae) with an analysis of their phylogenetic relationships. The Canadian Entomologist 123: 627-704.

9. Alarie, Y., and Maire, A. 1991. The dytiscid fauna of Québec Subarctic (Coleoptera: Dytiscidae).  The Coleopterists Bulletin 45(4): 350-357.

8. Alarie, Y. and Harper, P.P.  1990.  Primary setae and pores on the last abdominal segment and the urogomphi of larval Hydroporinae (Coleoptera: Dytiscidae) with notes on other dytiscid larvae. Canadian Journal of Zoology  68: 368-374.

7. Alarie, Y., Harper, P.P., and Maire, A.  1990.  Primary setae and pores on legs of larvae of Nearctic Hydroporinae (Coleoptera: Dytiscidae).  Quaestiones Entomologicae 26: 199-210.

6. Alarie, Y. Harper, P.P., and Roughley, R.E.  1990.  Description of the larvae of eleven Nearctic species of Hygrotus Stephens (Coleoptera: Dytiscidae: Hydroporinae) with an analysis of their phyletic relationships.  The Canadian Entomologist 122: 985-1035.

5. Alarie, Y. 1989.  The larvae of Laccornis Des Gozis 1914 (Coleoptera: Adephaga: Dytiscidae) with description of L. latens (Fall, 1937), and redescription of the mature larva of L. conoideus (LeConte, 1850).  Coleopterists Bulletin 43: 365-378.

4. Alarie, Y., Harper, P.P., and Maire, A. 1989. Rearing dytiscid beetles (Coleoptera: Dytiscidae).  Entomologica Basiliensia 13: 147-149.

3. Alarie, Y. and Leclair, R. Jr. 1988. Water beetles records from shallow pools in southern Québec (Coleoptera: Dytiscidae). The Coleopterists Bulletin  42: 353-358.

2. Bourassa, J.P., Alarie, Y., and Leclair, R., Jr. 1986. Distribution and habitat selection of dytiscid beetles in characteristic vegetal units of Southern Québec. Entomologica Basiliensia 11: 289-296.

1. Leclair, R. Jr., Y. Alarie, and Bourassa, J.P.  1986. Prey choice in larval Dytiscus harrisii Kirby and D. verticallis Say (Coleoptera: Dytiscidae). Entomologica Basiliensia 11: 337-342.